一名作家在作品未完成之时不幸撒手人寰，留下了作品中没有结局的王子（矢剃直树 配音）独自徘徊，王子和故事中的宿敌乌鸦来到了现实世界中，为了保护现实世界里的生灵，王子用自己的心封印了邪恶的乌鸦。 作家的鬼魂不甘心看到自己的故事就此停滞，他给了一只鸭子一条神奇的项链，鸭子因此而化身成为了雅喜儿（加藤奈奈绘 配音），她的使命就是找到王子因为封印乌鸦而四处散落的心之碎片。一次偶然中，法齐亚（樱井孝宏 配音）拾得了王子的心脏，将其取名为缪，害怕雅喜儿将缪诗夺走的法齐亚禁止两人见面，但缪诗反而因此而对雅喜儿充满了好奇。Ahiru (literally 'Duck' in Japanese), is a petite young girl in a junior high school's beginner ballet program. She's prone to stammering and clutziness. She has a huge crush on her sempai, Mute, a sad-seeming young man who doesn't speak much. Ahiru discovers that Mute is actually the prince out of a story written by the mysterious writer Drosselmeyer. When Drosselmeyer died, the prince and the evil crow he was battling in the story escaped. The prince defeated the crow, but only at the expense of shattering his own heart. Mute is a boy without feeling or understanding. Drosselmeyer, however, has somehow returned, and has offered Ahiru a chance to help Mute. She must become Princess Tutu, a magical ballerina, and help reclaim the pieces of her prince's heart. There is a catch, of course--Mute is being controlled by his roommate Fakia, also a dancer, who seems to want to keep the boy soulless and heartless. Drosselmeyer also told Ahiru that a certain dream she's been having is real. She really is just a little duck, and has been transformed into a girl only so that she can become Princess Tutu. If she starts stammering-- quacking like a duck-- she'll turn back into one.
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